Alionoctula Kruskop, Solovyeva & Kaznadzey, 2018. Zool. Stud. 57: 60
Vesperugo stenopterus Dobson, 1875
References:
Kruskop, S.V., E.N. Solovyeva, and A.D. Kaznadzey. 2018. Unusual Pipistrelle: Taxonomic Position of the Malayan Noctule (Pipistrellus stenopterus; Vespertilionidae; Chiroptera). Zoological Studies 57(60): 1-15. Read article.
Zhukova, S.S., A.P. Yuzefovich, V.S. Lebedev, and S.V. Kruskop. 2025. Reassessment of the Taxonomic Borders Within Pipistrellus (Chiroptera, Vespertilionidae, Pipistrellini) . Diversity 17(5): 1-24. Read article.
Alionoctula abramus (Temminck, 1838).
Mongr. Mamm. Tome 2: 232.
Japanese Pipistrelle
Alionoctula adamsi (Kitchener, Caputi & Jones, 1986).
Rec. West. Aust. Mus. 12: 463.
Adams' Pipistrelle
Alionoctula angulata (Peters, 1880).
Sitz. Ges. Naturf. Freunde p. 122.
Angulate Pipistrelle
Alionoctula babu (Thomas, 1915).
J. Bombay Nat. Hist. Soc. 24:30.
Himalayan Pipistrelle
Pipistrellus babu
Pakistan, Muree
Afghanistan through Pakistan, N India, Nepal, Bhutan, Myanmar, and into China
CITES - Not Listed (2023). IUCN - Least Concern under Pipistrellus javanicus (2019).
javanica species group. Formerly included in Pipistrellus; see Zhukova et al., 2025. Formerly included in javanica, but distinct; see Saikia et al. (2025). For earlier consideration of the placement of this species as a part of javanica see Corbet and Hill (1992), Kock (1996), and Bates and Harrison (1997), as well as Das (1990) and Sinha (1999).
References:
Saikia, U., R. Chakravarty, G. Csorba, M.A. Laskar, and M. Ruedi. 2025. Taxonomic reassessment of bats from the Western Himalayas, India and description of a new species of the Myotis frater complex (Mammalia, Chiroptera, Vespertilionidae). Zootaxa 5644(1): 1-78. Read article.
Thomas, O. 1915. Scientific results from the mammal survey. A. On pipistrels of the genera Pipistrellus and Scotozous. Journal of the Bombay Natural History Society 24(1): 29-34. Read article.
Alionoctula ceylonica (Kelaart, 1852).
Prodr. Faun. Zeylanica p. 22.
Kelaart's Pipistrelle
Alionoctula collina (Thomas, 1920).
Ann. Mag. Nat. Hist. ser. 9, 6: 533.
Greater Papuan Pipistrelle
Alionoctula coromandra (Gray, 1838).
Mag. Zool. Bot. 2: 498.
Indian Pipistrelle
Alionoctula dhofarensis (Benda, Reiter, Uhrin & Varadínova, 2016).
Acta Chiropterol. 18(2): 311.
Dhofar Pipistrelle
Alionoctula endoi (Imaizumi, 1959).
Bull. Natl. Sci. Mus. Tokyo 4: 363.
Endo's Pipistrelle
Alionoctula javanica (Gray, 1838).
Mag. Zool. Bot. 2: 498.
Javan Pipistrelle
Alionoctula minahassae (Meyer, 1899).
Abh. Zool. Anthrop.- Ethnology. Mus. Dresden 7(7): 14.
Minahassa Pipistrelle
Alionoctula murrayi (Andrews, 1900).
A Monograph of Christmas Island (Indian Ocean) p. 26.
Christmas Island Pipistrelle
Christmas Island (Indian Ocean)
Christmas Island (Indian Ocean)
CITES - Not Listed (2023). IUCN - Extinct (2017).
Formerly included in Pipistrellus; see Zhukova et al., 2025. Long recognized as a distinct species (see e.g., Chasen, 1940; Tate, 1942; Kitchener et al., 1986; Hill and Harrison, 1987), but synonymized with tenuis by Koopman (1973), a treatment followed by many subsequent authors (e.g., Corbet and Hill, 1992; Simmons, 2005). Morphologically and molecularly distinct from tenuis; see Helgen et al. (2009). Despite extensive efforts to locate additional individuals, the last sighting of this species occurred in August 2009 (Lumsden et al., 2021). Occasional strays of murrayi were observed on Cocos-Keeling Island; see Wood-Jones (1910).
References:
Chasen, F.N. 1940. A handlist of Malaysian mammals: a systematic list of the mammals of the Malay Peninsula, Sumatra, Borneo, and Java, including the adjacent small islands. Bulletin of the Raffles Museum (Singapore) 15: 1-209. Read article.
Corbet, G.B., and J.E. Hill 1992. Mammals of the Indomalayan Region. A Systematic Review. Oxford University Press, Oxford: 1-488. Not available online.
Helgen, K.M., K.N. Armstrong, J. Guzinski, R.A. How, and S.C. Donnellan. 2009. Taxonomic status of the Christmas Island Pipistrelle, Pipistrellus murrayi Andrews, 1900, as assessed by morphometric and molecular phylogenetic investigations of Indo-Australian Pipistrellus. Report to the Commonwealt: 26 pp. Not available online.
Hill, J.E., and D.L. Harrison. 1987. The baculum in the Vespertilioninae (Chiroptera: Vespertilionidae) with a systematic review, a synopsis of Pipistrellus and Eptesicus, and the description of a new genus and subgenus. Bulletin of the British Museum (Natural History), Zoology Series 52(7): 225-305. Read article.
Kitchener, D.J., N. Caputi, and B. Jones. 1986. Revision of Australo-Papuan Pipistrellus and Falsistrellus (Microchiroptera: Vespertilionidae). Records of the Western Australian Museum 12: 435-495. Read article.
Koopman, K.F. 1973. Systematics of Indo-Australian Pipistrellus. Periodicum Biologorum Zagreb 75: 113-116. Not available online.
Lumsden, L.F., P.A. Racey, and A.M. Hutson. 2021. Pipistrellus murrayi (errata version). The IUCN Red List of Threatened Species 2017: T136769A209549918. Read article.
Simmons, N.B. 2005. Chiroptera. Pages 312-529 In D.E. Wilson and D.M. Reeder (eds.) Mammal Species of the World: A Taxonomic and Geographic Reference, 3rd edition, Volume 1. Baltimore: Johns Hopkins University Press. Read chapter.
Tate, G.H.H. 1942. Review of the vespertilionine bats: with special attention to genera and species of the Archbold collections. Bulletin of the American Museum of Natural History 8(7): 1-297. Read article.
Wood-Jones, F. 1910. Coral and Atolls. Lovell Reeve & Co., London: 454 pp. Read book.
Zhukova, S.S., A.P. Yuzefovich, V.S. Lebedev, and S.V. Kruskop. 2025. Reassessment of the Taxonomic Borders Within Pipistrellus (Chiroptera, Vespertilionidae, Pipistrellini) . Diversity 17(5): 1-24. Read article.
Alionoctula papuana (Peters & Doria, 1881).
Ann. Mus. Stor. Nat. Genova 16: 696.
Lesser Papuan Pipistrelle
Alionoctula paterculus (Thomas, 1915).
J. Bombay Nat. Hist. Soc. 24: 32.
Mount Popa Pipistrelle
Alionoctula stenoptera (Dobson, 1875).
Proc. Zool. Soc. Lond. 1875: 470.
Narrow-winged Pipistrelle
Alionoctula sturdeei (Thomas, 1915).
Ann. Mag. Nat. Hist. ser. 8, 15: 230.
Sturdee's Pipistrelle
Japan, Bonin Isls, Hillsboro (= Hahajima) Isl
Bonin Isls (Japan)
CITES - Not Listed (2023). IUCN - Extinct (2020).
Formerly included in Pipistrellus; see Zhukova et al., 2025. Reviewed by Yoshiyuki (1989) and Görföl et al. (2020). Known from a single female "slightly immature" (Thomas, 1915: 231) specimen collected in the 1890s with an uncertain provenance (see Görföl et al., 2020), this species has not been reported since and is likely extinct.
References:
Görföl, T., D. Fukui, and G. Csorba. 2020. The taxonomic reassessment of a reportedly extinct bat, Pipistrellus sturdeei (Chiroptera: Vespertilionidae). Zootaxa 4755(1): 163-170. Read abstract.
Thomas, O. 1915. On the bats of the genera Nyctalus, Tylonycteris, and Pipistrellus. Annals and Magazine of Natural History ser. 8, 15(86): 225-232. Read article.
Yoshiyuki, M. 1989. A systematic study of the Japanese Chiroptera. National Science Museum Monographs 16: 1-242. Read abstract.
Zhukova, S.S., A.P. Yuzefovich, V.S. Lebedev, and S.V. Kruskop. 2025. Reassessment of the Taxonomic Borders Within Pipistrellus (Chiroptera, Vespertilionidae, Pipistrellini) . Diversity 17(5): 1-24. Read article.
Alionoctula tenuis (Temminck, 1840).
Monogr. Mamm. 2: 229.
Least Pipistrelle
Alionoctula wattsi (Kitchener, Caputi & Jones, 1986).
Rec. West. Aust. Mus. 12: 472.
Watts' Pipistrelle
Alionoctula westralis (Koopman, 1984).
Amer. Mus. Novit. 2778: 13.
Koopman's Pipistrelle
Includes all "eastern" pipistrelles (abramus, ceylonica, javanica, tenuis, coromandra, paterculus, and dhofarensis) and related forms (e.g., cf. coromandra) and tentatively all Pipistrellus species in Southern and East Asia and Australasia. Includes three species groups. The "javanicus species group includes the "eastern" species noted above. The "abramus" group includes only abramus itself. The "stenoptera" group also includes the single species stenoptera. Zhukova et al. suggest that adamsi, angulata, collina, papuana, and wattsi belong to the "javanica" species group. We have not allocated westralis, endoi, minhassae, murrayi, or sturdeei into any species groups within Alionoctula.